SAMENVATTING
SAMENVATTING Vooral na de conferentie van de Verenigde Naties in 1977 in Nairobi is het duidelijk geworden dat 'woestijnvorming' een probleem is dat voor alle aride gebieden over de gehele wereld geldt. De situatie in grote delen van Zuidwest Azie, waar in deze eeuw, met name in de laatste decennia, veranderingen zijn opgetreden in de wijze en intensiteit van het landgebruik, is een duidelijk voorbeeld. Vee1 van de veranderingen leiden op grote schaal tot een ernstige aantasting en uitputting van natuurlijke hulpbronnen, met name van fauna, vegetatie en bodem. Hierdoorwordt in toenemende mate land minder bruikbaar of ze1fs onbruikbaar. Als begrazing het algemeen aanvaarde 1andgebruik is, neemt de vegetatie a1s natuurlijke hu1pbron daarbij een sleute1positie in. Tegen deze achtergrond behandelt deze studie de al eeuwen1ang begraasde woestijngebieden van Irak, waarbij de nadruk ligt op de oeco1ogie en het gebruik van de vegetatie en op de mens als oeco1ogische faktor. Het doe1 van de studie is het 1everen van een bijdrage aan een verantwoord gebruik onder een gericht beheer van deze gebieden. Het veldwerk is uitgevoerd van april 1970 tot juni 1973 vanuit het Institute for Applied Research on Natural Resources, nabij Bagdad. Het onderzoeksgebied wordt gevormd door de Southern Desert, Western Desert en Lower Jezira (Fig. 1-3), tezamen ongeveer 240000 km 2 groot. Het plantendek bestaat er voor het overgrote deel uit lage, open struikvegetaties waartussen, na de winter- en voorjaarsregens, annuellen en bo1gewassen opb1oeien. Min of meer overeenkomstige vegetatietypen bedekken ongeveer 10-15% van het landoppervlak van de aarde; zij vormen een natuurlijke hulpbron van grote betekenis, vooral in verband met extensieve veeteelt. De onderzochte woestijngebieden vormen ongeveer 55% van het oppervlak van Irak, een land dat vooral bekend is door de vroegere culturen in en langs de Mesopotarnische vlakte. Olie vormt er nu de belangrijke b~on van inkomsten. Bodem, vegetatie en fauna van de woestijngebieden van Irak zijn, door de marginaal beschouwde gebruiksmogelijkheden, over het algemeen nog in geringe mate of in het geheel niet onderzocht. Enke1e k1eine gebiedjes vormen hierop een uitzondering. Aan de floristische en p1antengeografische aspecten van de vegetatie is relatief veel aandacht besteed, voora1 door verzamelaars tijdens studiereizen. Het in deze stu die onderzochte gebied he eft een gerniddelde jaarlijkse regenval varierend van ongeveer 50 mm tot 350 mm, met enorme verschillen tussen de jaren. Het grootste dee1 van de regen valt in enke1e hevige buien. De temperaturen dalen's winters soms tot bene den het vriespunt en lopen 's zomers op tot ongeveer SO°C. De jaarlijkse verdamping bedraagt ongeveer twee tot drie en een halve meter en de relatieve luchtvochtigheid kan in de zomer overdag dalen tot beneden tien. procent. Het gebied kan worden verdeeld in zeven fysiografische eenheden die zich vooral van elkaar onderscheiden door relief, het meer of rninder doorsneden zijn met wadis, zandig of stenig karakter en, daarmee samenhangend, bodemtype en hoogteligging (ongeveer 0-91 S meter boven gerniddeld zeeniveau). Deze 403
eenheden en de voorkomende bodemtypen worden kort besproken. Van groot belang voor de exploitatie van de vegetatie is de watervoorziening. Oorspronkelijk was men daarvoor aangewezen op enkele diepe gegraven putten, ondiepe putten veelal in of nabij wadis, enkele natuurlijke bronnen en (slechts van korte duur) verzameld regenwater dat aan het oppervlak bleef staan. Vanaf de jaren dertig zijn er putten geslagen. In de loop der jaren is het aantal daarvan uitgegroeid tot enkele honderden. Deze zijn soms tot honderden meters diep geboord en voorzien van pompen en opslagtanks. Deze verbetering in de watervoorziening en het gebruik van vrachtwagens en ander gemotoriseerd vervoer hebben vrijwel de gehele woestijn binnen het bereik van mens en vee gebracht. Na het in gebruik raken van vuurwapens en vooral met de invoering van snelle transportmogelijkheden werd er meedogenloos gejaagd, vooral op de grotere diersoorten. Oryx, struisvogel en andere diersoorten die hier eens in grote groepen voorkwamen zijn thans uitgeroeid evenaIs de grote roofdieren zoals de leeuw en het jachtluipaard. Nog slechts gazellen komen sporadisch voor. De mens moet al zeer lang geleden in dit gebied aanwezig zijn geweest, getuige de gevonden artefacten. Van een duidelijke invloed op de vegetatie kan echter pas sprake geworden zijn na het ontstaan van het nomadisme, toen Bedoeiimen vanuit het Arabisch schiereiland naar het Noorden trokken in het eerste en tweede millennium v. Chr., na het domesticeren van de dromedaris, de eenbultige kameel. Eeuwenlang is daarna de nomadische begrazing het landgebruik geweest voor de woestijngebieden. De Bedoeiimenstammen hadden hun eigen strikte regels voor het gebruiksrecht van bepaalde streken. Vooral de beperkingen in de drinkwatervoorziening en de vijandelijkheden tussen de stammen hielden de totale begrazingsdruk laag. Na de eerste wereldoorlog is dat beeld snel veranderd. De grenzen van het huidige Irak werden vastgelegd, er kwam leger- en politietoezicht, en met snelle transportmogelijkheden en zich steeds uitbreidende drinkwater-voorzieningen werd de woestijn 'opengelegd'. De kuddes kamelen en vooral schapen groeiden; voor geheel Irak nam het aantal schapen toe van 5 525 000 in 1938 tot 11 040 000 in 1965. Dit proces en de gevolgen ervan voor de vegetatie zijn nagegaan. Gebaseerd op een verkenning van het gebied, waarvoor van ongeveer 4000 km een globale besehrijving van terrein, bodem, vegetatie en landgebruik werd gemaakt, is, na een subjeetieve selektie, op 54 plaatsen een gedetailleerde vegetatie bemonstering uitgevoerd. In 50 kwadraten van 4 m2 per plaats werden diehtheid, hoogte, bedekking en (als totaalgetal) biomassa van de voorkomende overblijvende planten bepaald. Tevens werd de bodem bemonsterd op twee punten per plaats op de dieptes 0-30 cm en 30-60 em. De betrouwbaarheid en de bewerking van deze gegevens voor een c1assifieatie worden besproken. Bij de verwerking van de gegevens is een nieuwe kwantitatieve maat gebruikt. Daarbij is in een percentage (het gerniddelde van relatieve diehtheid, relatieve frequentie en relatieve bedekking) de bijdrage van de soort in de totaalopname uitgedrukt. Uiteindelijk werden 15 vegetatietypen en 14 socio-oeeologische groepen onder404
scheiden. De oecologische basis voor deze typen is met name gevonden in een klimaatsgradient, een verschil in bodemtextuur, en het landgebruik, waarbij vooral het ploegen van de bodem in het verleden meespeelt. De bedekkingspercentages liggen in de grootte orde van 1- 20% met slechts enkele hogere waarden. Het aantal soorten is laag. De autoecologie van de drie belangrijkste soorten, Haloxylon salicornicum, Rhanterium epapposum en Artemisia herba-alba, wordt behandeld, waarbij de nadruk ligt op de oecologie van de voortplanting. De eerste twee soorten vinden hun noordelijkste verspreidingsgrens in Irak. Het kiemingsproces en de ontwikkeling van de kiemplanten van de soorten illustreren hun aanpassing aan de extreme omstandigheden. Voor Rhanterium epapposum, een composiet van zandige bodems, wordt voor het eerst het kiemingsproces beschreven. In de gevallen bloemhoofdjes kiemen na de regens meerdere zaden terwijl ze in het hoofdje blijven zitten. De worteltjes dringen door de gemeenschappelijke bloembodem in het vochtige zand. Er is geen reden om aan te nemen dat door concurrentie over het algemeen slechts een kiemplant verder uit zou groeien. Uit een vergelijking van de strategie van de voortplanting wordt duidelijk hoe elk van de so orten aan een bepaald milieu is aangepast. In een apart deel worden het gebruik van de woestijngebieden door de mens en vee, in het bijzonder de vegetatie, en de gevolgen van dit gebruik, uitgewerkt. Begrazing door kamelen en schapen, thans vooral door schapen (enkele miljoenen), en in mindere mate geiten en ezels, heeft van oudsher plaatsgevonden, maar door de toegenomen begrazingsdruk is de vegetatie recentelijk sterk aangetast. De nieuw verzamelde en de reeds bekende gegevens over de voorkeur van de vee so orten voor de meest algemene plantensoorten zijn samengevat. Chemische analyse van plantenmateriaal toonde aan dat de bij voorkeur gegeten delen van de struiken een hoog ruw eiwit gehalte hebben en vaak rijk zijn aan mineralen; de gehaltes liggen zelfs hoger dan bij gekweekte voedergewassen. Het vochtgehalte varieert sterk over de seizoenen. In het voorjaar kan de vegetatie (inc1usief de annuellen) lOveel vocht bevatten dat extra drinkwater voor het vee overbodig wordt. Van geselecteerde soorten is de produktie over de seizoenen en, in een geval, voor twee jaren vergeleken. Door de ongeveer 4000 km verkenningstochten als transecten te analyseren en door gebruik te maken van de beschikbare produktiegetallen, is een globale schatting gemaakt van de totale produktie in de 240000 km 2 in 1971. Dit cijfer (2,91 x 109 kg drooggewicht) is vergeleken met de behoefte aan veevoer, afgeleid uit de 1971 - census gegevens (2,52 - 3,31 x 109 kg drooggewicht). Deze cijfers zijn voor het begin van de jaren zeventig goed met elkaar in overeenstemming. De produktie in de midden zestiger jaren, en waarschijnlijk ook daarvoor, moet echter veel hoger zijn geweest gezien de aantallen stuks vee, en de daarmee samenhangende behoefte aan voer, vermeld voor de 1965 census. Verdere veldgegevens, die een aanwijzing zijn voor de afgenomen produktie, worden besproken. Tevens blijkt de produktie per millimeter regen voor de overblijvende vegetatie slechts ongeveer een kilogram per hectare te bedragen 405
terwijl in de literatuur voor vergelijkbare gebieden in Noord Afrika drie tot vier kilogram worden vermeld. Het kappen van struiken voor brandstof (koken, verwarming) is een gebruik dat ondanks zijn omvang op zichzelf geen emstige gevolgen hoeft te hebben. Dat dit wei het geval is komt doordat het voortdurend plaatsvindt in dezelfde beperkte gebieden, rondom dorpen, putten, bronnen, etc., en bovendien omdat de eventuele hergroei weer aan be grazing blootstaat. Experimenten, waarin struiken van Haloxylon salicornicum en Rhanterium epapposum (herhaaldelijk) werden geknipt, tonen enerzijds, zolang het knippen niet te frequent plaatsvindt, de relatief grote weerstand van de eerste soort, maar tonen anderzijds de desastreuze gevolgen van het gebruik zoals dat door de bevolking, in combinatie met het vee, plaatsvindt. De meest direkte vemietiging van de natuurlijke vegetatie heeft plaatsgevonden doordat lager gelegen gebieden, die aflopend regen water ontvangen, mechanisch worden geploegd in (onverantwoorde) pogingen een graangewas te verbouwen. De meest produktieve vegetaties vallen hier onder en regeneratie vindt niet of nauwelijks plaats. Dit proces wordt besproken voor de noordelijke Jezira, het gebied met de hoogste regenval-verwachting, waar vrijwel geen natuurlijke vegetatie is overgebleven, en tevens voor de wadis en later gelegen terreinen in de woestijn in het zuiden. Na ploegen blijkt in veel gevallen een vegetatie op te komen waarin Peganum harmala en Artemisia scoparia domineren, twee soorten die voor begrazing of ander gebruik vrijwel geen waarde hebben. Waamemingen aan gebieden die een aantal jaren strikt beschermd zijn geweest of slechts licht begraasd tonen aan hoe hoog de bedekking, biomassa en produktie van bepaalde vegetaties kunnen worden onder dergelijke omstandigheden. Suksessie treedt wei op, maar blijkt voor de struikvegetaties een uiterst langzaam proces. Vergelijkingen van gebieden binnen en buiten afrasteringen geven een driemaal meer of nog hogere biomassa en bedekking voor de gebieden binnen de afrasteringen. Onder strikte bescherming blijkt de produktie van Haloxylon salicornicum terug te lopeno Een niet te intensieve begrazing stimuleert bij deze soort de groei. De gecombineerde en elkaar versterkende gevolgen van overbegrazing, kappen voor brandstof en ploegen, in diverse combinaties, tonen duidelijk aan hoe ver de aantasting van de vegetatie, en ook van de bodem door enorme erosie, al is voortgeschreden. Voor veel gebieden zal regeneratie moeilijk zijn of zelfs onmogelijk blijken. De verwachting is daarom dat, als de aantasting kan doorgaan in het huidige tempo, de eens zo produktieve woestijn-begrazingsgebieden van leak over enkele decennia zullen zijn omgezet in een kale, levenloze woestijn. Deze 'woestijnvorming' is geen proces dat zich over een zeker front in een bepaald tempo afspeelt. Het is veeleer een aantasting van binnen uit, waarbij de invloedssferen van de aangetaste gebieden elkaar tenslotte gaan raken, en waardoor geleidelijk de natuurlijke hulpbronnen van het gebied in zijn geheel worden aangetast en vernietigd. 406
In het 1aatste dee1 van de studie wordt een aanpak uitgewerkt die tot een beter beheer kan 1eiden. Een argument hiervoor kan ook gevonden worden in de economische betekenis van het gebied, in ve1erlei opzicht, maar voora1 voor de veetee1t. Deze betekenis is afzonderlijk uitgewerkt. Voor de periode 1953-1961 waarvan vo1doende gegevens beschikbaar waren, blijkt dat de woestijngebieden al1een al door de veeteelt 10-15% bijdroegen tot het totale aandee1 van de 1andbouwsector aan het nationaa1 inkomen. Tevens blijkt uit de sne1 stijgende v1eesprijzen, door de groeiende bevo1king en het toenemend inkomen, de toenemende vraag naar veeteeltprodukten. Om tot een verantwoord beheer te komen wordt een aanpak in drie fasen uitgewerkt waarin de 1andeva1uatie-procedure, zoa1s die onlangs door de F AO is voorgeste1d, een be1angrijke ro1 speelt. In de eerste fase wordt op basis van inventarisatie en k1einschalige kaarten van de be1angrijkste onderde1en van het land, zoals terreingeste1dheid, bodem, vegetatie, etc. voor het gehe1e gebied een globa1e geschiktheidsc1assificatie voor verschillend 1andgebruik uitgewerkt. Deze 1andgebruikstypen en de daarvoor in aanmerking te nemen faktoren worden afzonderlijk besproken. De overheid zou hieruit een keuze moeten maken en prioriteitsgebieden aangeven. In de vo1gende fase worden gese1ecteerde gebieden op een tienmaa1 grotere schaal gekarteerd. Er wordt dan ook een eva1uatie met een kosten/baten verwachting gemaakt, waarbij rekening wordt gehouden met de sociaa1-economische faktoren. Op basis hiervan moe ten in de derde fase concrete beheersmaatrege1en worden doorgevoerd waarvoor een sterke organisatie in het 1even geroepen zou moe ten worden. De ontwikkeling in de sociaal-economische sector zowe1 als veranderingen in de vegetatie en bodemgeste1dheid dienen dan nauwkeurig gevo1gd te worden waarbij een voortdurend overleg tussen de waarnemers, overheid, beheerders en gebruikers tens10tte een optimaa11andgebruik zou moeten garanderen. Enke1e aspecten van de kaarteringen, en het zo be1angnjke meten van de produktie van de vegetatie, worden afzonderlijk behande1d. Onder toepassing van moderne survey-technieken zouden de kaarteringen re1atief sne1 en efficient kunnen worden uitgevoerd. De bovengrondse biomassa en produktie van de struikvegetaties blijken met vrij grote betrouwbaarheid af te lei den uit de gemakkelijk te meten parameters diameters en hoogte van de individue1e struiken. Tenslotte worden de ontwikkelingen kort samengevat. Dezelfde technische ontwikkelingen die ge1eid hebben tot de enorme aantasting en degradatie van de woestijnbegrazingsgebieden zouden kunnen worden aangewend om tot een verantwoord beheer van de nog overgeb1even natuurlijke hu1pbronnen te komen. De methoden daarvoor zijn in grote lijnen voorhanden en de inkomsten uit olie-export p1aatsen Irak in een re1atief gunstige positie.
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INDEXES
AUTHOR INDEX Aart, R. van, 18 Abdulaziz, A.I., see AI-Ani, T.A. Aboona, S.H., see KhawaJa, A.K. Abu EI-Souod, S., 125 Abul-Fatih, H. (Hussain) A., 23, 34, 84, 121,141,161 Agnew, A.D.Q., 22, 31, 33, 34, 197, 314, 315,322,328,329,335,341 Ahmed, Adnan M., 67, 87, 96 Ahmed, Mohammed M.A., 118, 349, 352, 354,358 Ainsworth, W. esq., 300, 389 AI-Amari, Ahmed Shehab, 36 AI-Ani, Hikmat A., 33 AI-Ani, T. (Tariq) A., 7, 23, 34,43,79,121, 122, 132, 174, 185, 186,187,205,230, 235,236,237,240,241 see AI-Mufti, M.M. see Clor, M.A. see Habib, I.M. AI-Barazanchi, Jamal Nuri, see AI-Amari, Ahmed Shehab AI-Charchafchi, F.M.R., see AI-Ani, T.A. see Clor, M.A. AI-Dessoubi, F., see Kassir, S.M. Alexander, L.T., see Clawson, M. AI-Hadad, A., see Sykes, J.G. AI-Hashirni, M.H., 36, 107, 354 AI-Hilli, Majeed R., see Hilli Ali, K. (Khalid) T., 36, 107 see Fine, N.e. see Sarsam, N.H. Ali, Omar Amin, see Fine, N.e. Alikhan, J., see Kassir, S.M. AI-Izzi, H., see Hassawy, G.S. AI-Jaff, F., see Kassir, S.M. AI-Jibouri, L., 207 AI-Kasab, Nafi Nasser, 30, 37, 65, 77, 111, 112, 115, 293, 295, 297, 298, 299, 370 AI-Khateeb, M., 31 Allen, Shirley Walter, 349 Allouse, B.E., 100 Allred, B.W., 378 see Williams, Robert E. AI-Morany, K., see Sykes, J.G. AI-Mufti, M.M., 7, 34, 35,43, 171, 180, 187, 204, 207, 208, 213, 253, 258, 260, 263,265,270,313,314,328,330 see AI-Ani, T.A. see Habib, I.M. AI-Nouri, H., see Sykes, J.G. AI-Oubaidi, Khalid S., see Jones, Richard G. AI-Rawi, A. (Ali), 31
see Townsend, e.C. AI-Roumi, Hikmat A., 349, 354 AI-Sabi, Ismail M., see Jones, Richard G. Altaie, F. (Flayeh) H., 76, 77, 78, 79, 82, 85,366 AI-Tawil, B.H., see Habib, I.M. AI-Tikriti, H., 31 AI-Uthman, Hilmi Sabir, 104 AI-Zubaidi, A., see Chaudri, 1.1. Anderson, C.A., 35, 270, 313 Asadi, R.F., see Khawaja, A.K. see Labib, A.1. Ashraf, A., 363 Aucher Eloy, P.M.B., 30 Babayev, A., see Kharin, N.G. Baig, M. Shabbir, 261, 364, 373, 375 Bakr,Z.Y., 169,185, 186, 187, 188 Bannink, J.F., 125 Barbour, Michael G., 186, 187, 188, 190,204 Barclay, A.S., 14 Barnes, R.F., 232 Bashir, N., see Kassir, S.M. Batanouny, K.H., 24, 33, 82,121,125,132, 133,141,145,151,156,173,174,301, 310 Bazilevich, N.I., 12, 13 see Rodin, L.E. Beard, John S., 131 Beaudelaire, J.P., 249 Beaumont, Peter, 16, 18 Beek, K.J., 363, 364, 368, 370, 371, 373 Beek, M.A., 18 Bell, Gertrude, 112 Bennema, J., 363 Blair, Robert M., see Short, Henry L. Blair Rains, A., 230, 231, 241 Blaisdell, James P., see McKell, Cyrus M. Blake, Gerald H., see Beaumont, Peter Blakelock, R.A., 8, 12,29,30 Bodenheimer, F.S., 32 Boissier, P.E., 31 Bor, N.L., 23,43, 159,228,229,322 Bormann, F.H., 124 Bornmiiller, J., 30 Box, Thadis W., see Stoddart, Laurence A. Braidwood, R.J., 109 Braun-Blanquet, J., 33, 132, 134, 136, 137, 140, 141 Brice, W.e., 16 Brichambaut, G. Perrin de, 22,36,293,295, 299,303 Brinkman, R., 363 Brown, Dorothy, 45, 106, 124, 249, 261, 378 431
Bruguiere, J.G., 30 Bryan, H.M., 35 Burdon, David J., 99 Buringh, P., 15, 16, 17, 18, 20, 30, 53, 74, 76,77,79,84,85,108,148,296,366 Burkart, Leonard, see Short, Henry L. Buschbom, U., see Kappen, L. Butzer, K. (Karl) W., 108, 109
Falcon, N.L., 17 Faraj, Mohammed, see Fine, N.e. Fine, N.C., 36 Firmin, R. 274 Fisher, W.B., 15 Floret, C. (Christian), 259, 260, 379 Flux, John, E.C., 197 Fosberg, F.R., 131 Freitag, H., 171, 269
Cain, S.A., 125 Castro, G.M. de 0., 125 Chakravarti, H.L., 31 Chapline, W.R., 14 Chapman, G.W., 21, 22 Chapman, R.A., 383 Chapman, V.1., 23 Chaudri, LL, 23, 34 Cheesman, R.E., 100, 102 Chesney, F.R., 30 Clawson, M., 5,16,18,349 Clifford, H.T., 137 Clor, M.A., 43, 183, 185, 186, 188, 189, 204,205,211 Condon, R.W., 374 see Moore, R.M. Cook,C. Wayne, 14, 230,238,240 Cooke, Ronald U., 74 Crampton, E.W., 230, 231
Gangopadhyahya, M., 53, 293 Gehrke, Ulrich, 113 Georg, P.V., 100,101,105 Gilbert, R.O., see Uresk, D.W. Gillett, J.B., 34, 165, 226, 228, 229,276, 303,304 . Gils, H.A.M.J. van, see Zonneveld, I.S. Gimingham, C.H., 250 Girgis, W.A., 131, 145 Goldman, Bram J., see McGinnies, William G. Good, R., 174 Goodall, D.W., 134, 137 Goodin, Joe R., see McKell, Cyrus M. Goosen, D.1., 377 Gounot, M., 134 Gounot, P., 6 Green, Christine R., 9 Greig-Smith, P., 44,124,125 Grigorshenko, LA., see Kharin, N.G. Grimsdell, J.J.R., 378 Gronovius, J.F., 30 Guest, E. (Evan) R., 8, 12, 14, 21, 22, 23, 24, 29,30, 31, 32, 33,34,43,53,60,65, 67,77,94,101,121,126,132,141,147, 150, 151, 153, 154, 155,157,158,160, 161, 165, 169, 178, 226, 228, 270, 274, 275,276,301,304,330 see Townsend, C.C.
Dakshini, K.M.M., 11 Dapper, 0., 30 Davis,P.H., 171, 174 Denio, Reginald M., see Williams, Robert E. Dennis, P.E., 53 Dickson, V., 110, 173, 174, 182, 197 Dietz, Donald, R., 14 Dijk, A. van, 377 Diskin, S., 251, 255 Doughti, Charles M., 110 Dougrameji, 1.,76, 150, 178 Draz, Omar, 371 Dregne, Harold E., 74, 75, 145 Dudley Stamp, L., 7, 349 Dijksterhuis, E.1., 133 Dzerdzeevski, B.L., 9 Eig, A., 30, 32, 161, 165 Elfyn Hughes, R., 127, 128,250 Ellenberg, Heinz, 45, 121, 122, 124, 125, 131,135 Emberger, L., 9, 36 Engel, e.G., 70 Evenari, M., see Kappen, L Everist, Selwyn L., 11 432
Habib, LM., 23, 34, 82 see AI-Ani, Tariq A. Hadac, E. (Emil), 22, 29, 31 Haddad, Sharif Y., 33 Haines, R.W. (Wheeler), 33, 34, 167, 169, 182,183,212,335,341 Hammouda, M.A., 169, 185, 186, 187,188 Handel-Mazzetti, H.F. von, 21, 22,30,31, 304 Hanna, A.Y., see Chaudri, 1.1. Harris, Grant A., 350 Harris, L.E., 230, 231 Harris, S.A., 23, 34 Harrison, D.L., 99, 100, 101, 102,103 Hasan, M.S., IH, 115 Haseeb, K., 106, 107, 352, 353, 354, 355,
356,357 Hassan, A. Hassan, 58 Hassawy, G.S., 23, 31 Hatt, Robert T., 99, 100, 101, 102, 105 Heady, Harold F., 219, 260, 265, 270, 349 Hielkema, J., 377 Hilli, (AI-) M. (Majeed) R., 24, 33,34,121, 132,133,141,145,147,150,151,156 Hills, E.S., 5 Hitti, Philip K., 392 Holt, A.L., 113 Hooker, J. (Joseph) D. (Dalton), 170, 171 Horwitz, W., 47 Hoste, C.H., 37, 106, 261, 266 Houerou, H.N. Ie, 11,37,106,132,111, 174, 259, 261, 263,265, 266, 270, 273 Hutchings, Selar, S., 379 Huzayyin, S., 111 Hyder, D.N., 12 Ibrahim, K.M., 29, 35, 36, 349 Jaccard, P., 136 Jackson, M.L., 47,48 Jacob, Georg, 390 Jamali, M.FJ., 111 Jawad, Maeda S., 230, 235, 237, 240 Johnson, Douglas L., 47,109 Johnson, Kendall L., see Morris, Meredith J. Jones, L., 18 Jones, Quentin, 14 Jones, Klcnard G., 354 Kappen, L., 171,212 Kareem, H.A., see Chaudri, l.l. Kassas, M. 131, 145 Kassir, S.M., 354 ~ul, R.N., 5,7,14,30,34,76,121,122, 150, 169, 171, 178, 180,204,207,208, 213, 253, 258, 260, 262, 263, 265, 270, 299, 314, 328, 329, 349, 354, 370, 379, 392 see AI-Ani, T.A. see AI-Mufti, M.M. Kellersohn, Heinrich, 36 Kernick, M.D., 37, 151,152,155,156,169, 173, 174, 178, 179, 209, 226, 228, 229, 260,261,265,270,274,349 Kershaw, Kenneth A., 125 Keulemans, N.C., see AI-Amari, Ahmed Shehab Kevork, Ohannes K., 104 Kharin, N.G., 379 Khawaja, A.K., 230, 233, 236, 241 Kinsinger, Floyd E., 127, 379 Kipling, K., see Tueller, P.T.
Kittani, H., 22 Knetsch, G., 6 Koppen, W., 9 Koster, Klaas, see Jones, Richard G. Kotchy, T., 30 Kozlowski, T.T., 243, 244 Krader,Lawrence,110 Kraft, Walter, 67, 87, 96 Kreeb, Karlheinz, 29 Krochmal, Arnold, 14 Kuhn, Gustav, 113 Labib, A.I., 230 Landsberg, H.H., see Clawson, M. Lange, O.L., see Kappen, L. Larfrere, L., 11 Larson, Gary E., 103 Lawless, R.I., 18, 114,115 Lawrence, T.E., 390 Lebon, J.H.G., 18,37,115 Lees, G.M., 17 Leigh, J.H., see Moore, R.M. Lewis, James K., 7, 350 Leys, H.N., see Bannink, J.F. Lieth, H. (Helmuth), 12, 13,65 Long,G.A.,35,36, 178, 270,314 Longrigg, S.H., 114, 115, 118 Lorain, G., see Tueller, P.T. Maarel, Eddy van der, 121, 125, 132, 140, 141 Mac Fadyen, W.A., 53, 87, 88, 89, 90, 92,94 Mahdi, Nuri, 100, 101, 105 Mahmood, Ghazi, 36 Maizels, Judith K., 5, 269, 340, 344, 349, 391 Malcolm R.T., see Post, W.S. Martonne, E. de, 9, 11,65 Mashkour, Mustafa, 58 Mason, Lamar R., 379 Matti, S.A., see Khawaja, A.K. see Labib, A.I. McFetridge, D.G., see Kassir, S.M. McGinnies, William G., 5,11,29,30 McKell, Cyrus M., 12 McLeroy, G.B., see Sarsam, N.H. Meigs, P., 9, 10, 11, 13 Michaux, A., 30 Michener, C.D., 136 Miller, G.R., 250 Milner, C., 127, 128, 249 Miroshnichenko, Yu.M., see Rodin, L.E. Mitchell, Raoul C., 67 Mobayen, S., 151, 174 433
Moen, Aaron N., 230, 231 Mokhtar, K.M., see Khawaja, A.K. Monakhov, M.A., 33,121 Mooney, H.F., 21 Moore, R.M., 7, 11 Morris, Meredith, 378 Mountfort, Guy, 100,104 Mueggler, Walter F., 371 Mueller-Dombois, Dieter, 45,121,122,124, 125,131,135 Musil, Alois, 110 Nabelek, Fr., 31 Nasserula, Sabah M., 34, 121, 141 Neal, Donald L., see Morris, Meredith J. Neale, P.E., 36 Noble, A.H., 87 Novikoff, G., 266, 273 Noy-Meir, Imanuel, 213, 253, 255, 266 Oedekoven, K.H., 22 Olivier, G.A., 30 'Oraim, Abdul Jabbar, 115 Orshan, G., 171, 212, 251, 255 . see Tadmor, N.H. Ouda, N., see AI-Ani, T.A. Paulsen, Harold, A. Jr., see Williams, Robert E. Paylore, Patricia, see McGinnies, William G. Pearse, C. Kenneth, 349 Pechanec, J.F., 350 Peppelenbosch, P.G.N., 110 Perry, R.A., 7,44,45,124,390,391 Petrov, M., 11, 270 Pitman, C.R.S., 100 Po1unin, N., 31 Pontanier, R., 259 Poore, M.E.D., 145, 173, 174, 178, 180, 304,349 Post, G.E., 43 Post, W.S., 36 see Al-Amari, Ahmed Shehab Poulton, C., 377 Preston, R.L., 230 Quezel, P. (Pierre), 170, 173, 174 Rasw:m. Carl R., I! 0, 112, 118 Rauwolff, L., 30 Rawitz, E., see Tadmor, N.H. Rechinger, K.H., 31, 43, 44,147,169,171 Regel, C. von, 31, 32 Reitan, Clayton H., 9 Rice, E.L., 135 434
Richard, W.H., see Uresk, D.W. Richards, L.A., 48 Risser, P.G., 135 Robertson, V.C., 145, 173, 174, 178, 180, 304,349 Robinette, W. Leslie, 14 Rodin, L.Y., 171, 253, 255 see Bazilevich, N.1. Roux, Georges, 30, 109 Rozov, N.N., see Bazilevich, N.1. Salek, K.M., see Labib, A.I. Sankary, Mohamed Nazir, 186, 187, 188, 190,204 Santa, S., 173 Sarsam, N.H., 36, 107 Schimper, A.F.W., 11 Schmidt-Nielsen, K., 243 Schulze, E.D., see Kappen, L. Schumacher, F.X., 383 Schwan, H.E., 35, 270 Serkahia, S.Y., 31, 36, 151, 165, 222, 223, 226,228 Sestini, l' Abbe, 30 Shantz, H.L., 11, 12, 13 Sharif, C.M., 243 Sharp, R.P., 70 Sheikh, M.Y., 33, 121, 141, 145 Shimwell, David W., 131 Short, Henry L., 14 Shreve, F., 145 Skouri, M., see Novikoff, G. Sluijs, A.A. van der, 30, 87, 97, 370 Smith, Arthur D., see Stoddart, Laurence A. Smyth, A.J., 363 Sobolev, L.N., 125 Soest, P.J. van, 232 Sokal, R.R., 136 Solecki, Ralph S., 108,109 Sorensen, T.A., 136 Soriano, Alberto, 11 Sprague, Howard S., 371 Springfield, H.W., 21, 33, 35, 36, 351, 352,358 Srivastava, A., 29 Stoddart, Laurence A., 230, 231, 243,261, 373,389,390,391 Stoops, G., see Altaie, Flayeh Strickler, Gerald S., 379 Suliman, 1., see Sykes, J.G. Sweet, Louise E., 118 Sykes, J.G., 354 Sys, H.C., see Altaie, Flayeh Tiickholm, Vivi, 43,169,173,174
Tadmor, N.H., 145 Takessian, B.A., see Habib, I.M. Tammimi, S.A., see Hassawy, G.S. Tammo, K. (Khairi), H. (Habib), see Ai-Amari, Ahmed Shehab see Post, W.S. Texier, C., 30 Thalen, D.C.P., 7, 30, 34, 35, 41,43,45, 79, 103, 121, 122, 169, 258, 260, 262, 263, 270, 313, 314, 330,349,354,366, 370,374,377,379,386,392 see Ai-Ani, T.A. see AI-Mufti, M.M. see Dijk, A. van see Zonneveld, I.S. Thornthwaite, C.W., 9, 11 Tigner, James R., 103 Tome, Rajaa Y., see Jones, Richard G. Townsend, C.C., 31, 43, 169 Tregubov, V., 151, 174 Tueiler, P.T., 378 Tukhaikh, Jabbar Kadhem, see Ai-Amari, Ahmed Shehab
Wagner, F.H., see Novikoff, G. Wagstaff, J. Malcolm, see Beaumont, Peter Wallen, e.e., 9, 293, 295, 299, 303 Walter, H., 65 Walton, K., 9 Warren, Andrew, 5, 74,269,340,344,349, 391 Weinert, Erich, 121, 141, 147, 150, 151, 156 Werger, M.J.A., 125, 133, 140, 162 West, Neil E., 243 Westhoff, Victor, 121, 125, 132, 140, 141 White, Gilbert F., 5 Whittaker, R. (Robert) H., 12, 13, 125, 130, 131,379 Whyte, R.O., 14, 108 Wierenga, T., 379 Wilkie, e., see Tueiler, P.T. Williams, Robert E., 349 Williams, W.F., 134, 137 Winkworth, R.E., 7, 30,41,270 Wrrth,Eugen, 16, 18,30,37,293, 299,306 Woodweil, G.M., 379
Ulrich, A., 47 Uresk, D.W., 379 Utekhin, V.D., 125 Uvarov, B.P., 32
Yacoub, Salah M., 115, 116 Yahia, H.M., 77, 84, 85,147 Yossef, S.H., see Labib, A.1.
Vailentine, John F., 7. 99 Van Dersal, William R., 14 Veen, J.P.H. van der, 36, 37,165,171,259, 265,270,273,274,310,349,354 Verstappen, H.Th., 377 Vesey Fitzgerald, Desmond Foster, 155, 169,173,174,178,179,182,195,210 Vink, A.P.A., 364 VOllte, Caesar Czn., 108
Zaki, M.A.F., 301, 310 Zand, G., 171, 212 Zohary, M., 21, 22, 23, 29, 30, 32, 65, 74, 75, 77, 121, 126, 132, 133, 141, 147, 150, 151, 156, 158, 165,167,169,171, 173,174,180,276 Zonneveld, I.S., 131, 363, 371, 377, 392 see Bannink, J.F. see Dijk, A. van
435
GENERAL INDEX Figures in italics refer to information in tables or figures. Abu Dhabi, 173, 174 Abu Dibbis, 23, 71, 79, 84, 104, 108, 150, 161,275,288,382,383 air humidity, 63, 64 air temperature, 64 soil temperature, 58 Abu Ghraib, 6, 127, 197 Abu Kemal, 73 Acacia iraquensis, 274 Acantholimon, 21,270 accessibility, 87, 176, 371 Achillea, 24,229,304,308 A. conferta, 143, 161,301,302,331 A. fragrantissima, 36, 14i, 143, 148,155, 158,160,179,222,226,229,233,234, 235, 236, 240,248, 277,307, 309,327, 328,329,330,338 Achillea fragrantissima - Artemisa herba· alba type, 160-161 Acinomyx jubatus Schreber, 102, 117 adaptive characteristics (of key species), 209-214 Adhaim river, 24, 276 Adonis dentata, 32 Aellenia subaphylla, 143 Aeluropus lagopoi"des, 23,141,143, 147, 161,228,248 Aeluropus littoralis, 23 aerial photography (interpretation), 43, 376,377,378 infra·red false colour, 378 large scale, 378, 386 aesthetic value, 14,350 afforestation, 97 Afghanistan, 171, 174,269 afterripening, 204, 211 Agathophora alopecuroides, 151, 315 agro-climatic regions (of Iraq), 351, 352 'ahrash',22 Aidaha, 42, 116 Ain Hamud, 87 Al Barit (AI Bereed), 94, 302 Al Dahnaii, 111 Aleppo, 30 alfalfa, 88, 241 Algeria, 173, 253, 259, 263, 270 Alhagi mannifera (A. maurorumj, 24, 137, 143, 161,301,302,331 Al Hajara Stony Plain, 68,69, 70,71,82, 84
Al Hamad Plain, 68, 71 Al Haniya Ridge, 70 Al Jauf, 174, 179 Alopecurus, 23 Al Qaim, 77,84 Amara, 22 Ana, 30,42, 53,100, 116 evaporation, 66 rainfall,59 Anabasis articulata, 212,255 Anabasis setifera, 104,143,151,222,226, 315,315,319,321,326 Anaiza, 174 Andrachne telephioides, 137, 143, 330, 331 Aneza (tribe), 112,296 animal unit, 106, 108,258, 261,357,358, 374 annuals (annual species, ephemeral species), 24,65,126,180,221,244,249,315,335 contribution to forage, 255-257, 255, 256-257 moisture content, 249, 250 in vegetation classification, 132-134 in vegetation types, 150-162 Ansab,42, 92, 116,275 Anvil/ea garcini, 143, 235,236, 240,315, 321 Aqarquf, Hor of, 111 aquifer, 96 Arabia, 100, 110,111,114,117,173,174, 178 see Saudi Arabia Arabian Gulf, 15, 17,54,63, 174 Arabian Plateau, 15 Arfajiya, 165 arid areas (general) delineation, 9-11 general characterizarion, 8 major plant formations, 13 world distribution, 10 Aristida obtusa, 36 Aristida plumosa, 36,100,125,315,321, 322 see Stipagrostis plumosa Armenian·Anatolian mountain region, 15 Arnebia decumbens, 250 Ar Rahab, 68,71,73,178 Artemisia, 270-304 A. absinthium, 300 A, fragrans, 300 A, herba·alba, 24, 32, 34, 36, 140,141, 143, 147,148,157,158,159,159,160, 160, 162, 165-214 (see specification below), 255, 265, 304, 314, 328, 338, 437
342,343 adaptive characteristics, establishment, 211, 212 maintenance, 212, 213 physiological, 212 reproduction, 209, 210, 211 chemical analysis and nutritive value, 233-240 description, 170-171 distribution, edaphic conditions, 177, 179-180,180 Iraq, 174-176, 175 general, 171-174 estimating biomass and production, 378-385 forage preference, 222, 223, 224, 225, 229, 229 forage production, 251,252, 253,260 forage value, 226 fuel, 277 germination, afterripening 204, 211 hydration and dehydration, 205, 205 process, 204 temperature, 205 grazing impact, 263, 263,264 (see Rutba exclosure) moisture content. 248, 249 phenology 181-182,181 protection, 328 (see Rutba exc1osure) rootsysteem, 213 seedling development, 205,206, 206, 207, 207,208,209 seed production and dispersal,204, 211 uprooting for cultivation, 294 Artemisia herba-alba - Helianthum lippii type, 158 Artemisia herba-alba- Salsola spp. type, 158-159 Artemisia scoparia, 24, 36, 73, 126, 140, 141, 147,148,156,157,158,159,160, 221,226,229,236,240,248,277,300, 301, 301,302,308,309,328,331,338, 343, Artemisia scoparia - Peganum harmala type, 161 artifacts, 108, 389 ash (minerals, content in forage), 230-231, 232,233,235, 236, 237,240,241,242, 372 Ash Shihiyat, 165 As Salman, 36,42, 53,70, 90, 102, 116, 271,275,275,314,341 rainfall, 60 Assyria, 30, 100 438
Astragalus, 21,174 A. spinosus, 33, 36, 141, 143, 153, 156, 157, 158,167,220,221,223,227,229, 229,248,270,328,338,339 Athaman, 43,276,309,309 Atriplex, 143, 236 A. leucaciada, 137,160,236,237 A. polycarpa, 187 Australia, 11, 12, 374,390,391 Avena, 23 A. barbata, 327 Baaj,42, 161,296,297,302 Baghdad, 6, 15, 18, 24, 30, 31, 42, 54, 101, 111, 112, 121, 145, 176, 270, 354,376 air humidity, 63, 64 air temperature, 56 evaporation, 65, 66 radiation, 55 rainfall, 59,60 soil temperature, 57 sunshine, 55 Bahrein, 173, 174 Baiji (Badji), 42, 76, 101 rainfall, 59 sand dunes, 34, 75-76,121,150 Balad Sinjar, see Sinjar Baluchistan, 171, 364 'barbak', 76, 88, 89 Barguth (depression), 74 Basra(h), 15, 18,22, 32, 42, 53, 54, 79, 111,151,176,198,276 air humidity, 63, 64 air temperature, 55 evaporation, 65,66 rainfall, 59, 60, 63 snowfall, 63 sunshine, 55 Bedouin(s), 88, 90, 95, 102, 104, 221, 223,243,271,272,273, 276,277,286, 296,297,335,340,349,390-391 changing population, 114-115 origin, 109-110 settling, 111-113, 116 sources of income, 113 Bienertia cycioptera, 34,143 birds, 103-105 Boara (Bouara) depression (salt lake), 23, 74,84,86 Boghars (depression), 74 Botswana, 43,366 boundaries (ofIraq), 15, 113-114, 118 Braun Blanquet approach, 132-134, 137, 140, 141 browse (browsing), 219, 374
see note page 219 buffalo (water-), 105,106,352-356 Busaiya (Bsaia), 35,42, 53,68,70,81, 94, 116, 151,152,153, 155,165,176, 178,195,201,271,274,275,276,277, 314,341 rainfall, 60 bustard, 104, 117 calcium (in forage), 231,232,235,236
Calendula aegyptiaca, 250 Calligonum comosum, 72,141,143, 150,
151, 180,227, 235,236, 240,276,277
Calligonum comosum - Cornulaca leucacantha type, 150-151
camel(s}, 23, 99, 105, 107, 111, 112, 113, 370 contribution to national income, 352359 numbers, 106 see grazing Canis aureus L., 102 Canis lupus L., 102 Capparis spinosa, 308, 310 caracal, 102, 103 Caracal caracal schmitzi Matschi, 102, 117 caravans, 113,301 carbohydrates, 230-231, 232, 240, 241, 242 Carex stenophylla, 24,32,161,222,327, 330,331 carrying capacity, 36 see grazing capacity Carthamus oxycanthus, 329 cattle, 99, 105, 106, 107, 112,352-356 census, agricultural, 107 human population, 18, 115 livestock population, 106, 107-108 Central Asia, 11, 270 charcoal,269, 270,276,350 cheetah, 102, 103, 117, 389 chemical analysis, plant material, 47 soils, 47-48 water, 97-99 see chemical composition chemical composition (of forage), 230243,236 differences in shrub species, 232-237 proximate analysis, 230-232 seasonal variation, 237-239 shrubs compared to other feeds, 240-243 chert, 70 Chlamydotis undulata Jacquin, 104-117
Citrullus colocynthis, 83, 221, 229, 329, 373 classification of vegetation, 130-142 characteristics, 130-134 dendrograms, 134, 137, 138, 139, 140, 141 guiding principles, 130-131 similarity indices, 135-137, 138, 139 syntaxonomy,141-142 climate, arid zone classification, 9-10 desert areas ofIraq, 53-67 factor for community types, 148-150 factor in land evaluation, 369-371 fluctuations, 70, 108 Iraq (general), 17 subdivision study area, 65-66 clustering (in vegetation classifications), 134,137 community type(s}, see vegetation types Cornulaca, 126, 150, 153,236,248, 301, 302, 321 C. leucacantha, 141, 143, 147, 148, 150 C. monacantha, 143,315 Cousinia spp., 143, 270,321 Crepis, 300 Cressa cretica, 23, 79, 161 crude fibre, see fibre crude protein, see protein Cynodon spp., 329 C. dactylon, 329 Cyperus, 100,152 C. conglomeratus, 100, 141, 143, 151, 152,152,248 Dahama,179 Darb Zubaida, 89 dates (date palms), 22, 88, 358 'decreasers', 133 dendrogram, see ciassfication of vegetation desert (definition), 8 desertification,S, 339-344, 349 see fuel (cutting) see grazing see ploughing desert monitor, 104 desert pavement, 74-75, 82, 179, 344 desert varnish, 70, 75 Dhahran, 173 Dibdibba (Plain) (area), 67,68,68, 69, 70, 71,76,79,97,154 digestibility, 230, 231,240,242 Diwaniya, 42 air humidity, 64 evaporation, 66 439
rainfall 59, 60, 63 snowfall, 63 Diyala river, 24 Diyala Liwa, 111 Dociostaurus maroccanus, 32 domestication, 17, 109, 350 dominance types (in vegetation classification), 131-132 donkeys, 105,106,270,271,276,352 Dulaim (tribe), 296, 300 dunes, see sanddunes see Euphrates sand-belt dung, 14,271,352 East Atrica. 378 EC (electrical conductivity) in soils, 48, 81,127,145,147,314 economic evaluation, 350-359 Egypt, 151, 173, 174, 185, 212, 301, Egyptian Desert, 131, 145 electrical conductivity, see EC Ephedra alata, 83, 104, 140, 141, 143, 147,150,153,153,155,155,156,158, 221, 223,227, 229,236,248, 270,277, 279, Ephedra alata - Rhanterium epapposum type, 152-153 Ephedra alata - Haloxylon salicornicum type, 155-156 ephemerals (ephemeral species), see annuals Equus hemionus hemippus Geoffroy, 100 Erodium sp., 331 E. cicutarium, 32,250, 327 E. deserti, 327 erosion, 17, 71, 74, 133,274,296,307, 309, 335,337, 338,340,341,342-343, 373,375 Eucalyptus, 275 Euphrates (river), River Euphrates, 15, 16, 17,30,67,70,71,73,77,87,88,110, 116,117,293,296,391 Euphrates sanddune- belt, 68, 68, 71, 72, 79,82,276 soil profile, 81 vegetation, 150-151 evaluation of range resources, 350-359 evaporation, 17, 18, 64, 66, 84, 145 exclosures (fenced areas), 34, 36, 212, 255,304,307,313-332,335,341 see Ha tra ex closure see Khedr-el-Mai ex closure see Rutba exclosure see Shabicha exclosure Fagonia bruguieri, 315,315
440
Faidhat al-Jeez, 157, 221, 279, 287, 309,338,339 Faidhat al-Mesad, 326 Fakka (exclosure), 314 Falco cherrug Gray, 104 falcons, 104 Falluja, 24,42, 175 Falluja Desert, 24, 33 fats (in forage), (Ether Extract), 230-231, 232,235,236,240,241,242 fauna, 99-108,117,369 Felis chaus Giildenstaedt, 102, 103 fibre, 230-231,232,234,235,236,237, 238, 239, 239-240, 241, 242 Filago spathulata, 32 firewood, see fuel floristics, in classification of vegetation, 131-134 species identification, 43-44 fodder crops, 36,241,241,295,351,370 Foothills zone, 16, 17,18,32 forage (production), 249-266, 328, 350 between year variability, 253-254,252, 372 contribution of annuals, 255, 255-257, 256-257 demand and supply, 257 - 266 estimation from plant attributes, 378386 exclosures, 318 land evaluation, 372 seasonal aspects, 251-253, 252, 372 see grazing forest, 19, 21-22, 25 foxes, 102 frost, 17 fuel (cutting), 14, 22, 23, 127, 148, 149, 269-289,269,277,278,279,288,295, 308,309,318-319,335-344,350,391 experimental fuel cutting, 41, 272, 278289,336,337 extent of the cutting, 272-274 fuel requirement, 272-273 fuel value of different vagetation types, 150-162 species involved, 274-277 use of the fuel, 270-271 Gagea reticulata, 32,154 game (animals), 100, 102, 350 see under the species Garrhadiolus hedypnois, 32 gas, 14, 392 Gazella spp., 100, 117 G. dorcas, 101,102
G. gazella, 101 G. subguttorosa, 101,102 gazelles, 100, 101,102,117,389 geological structure (of the study area), 67 gerbil, 103 Gerbillus spp., 103 G. dasyurus mesopotamiae Harrison, 103 germination, 209, 210, 211,335 Achillea fragrantissima, 330 Artemisia herba~alba, 204~205 Haloxylon salicornicum, 185~189 Rhanterium epapposum, 195, 196, 197~200
Ghurfa Desert, 24,33,121,132,133,145 goat~23,99, 105,106,109,370 contribution to national income, 352359 numbers, 106 see grazing granite, 178 grazing, 12,34,35,36,94,112,127,133, 148, 149, 208, 219~266, 270, 283, 289, 296,299, 300,308,319,322,327,329, 333,335~344,369,370,373
value of different vegetation types, 150-162 grazing capacity, 219, 257-266, 373, 374~375, 391 grazing pressure, 257-266, 326, 329.330 Greater Zab (river), 299 Great Nafud, 179 groundwater, 84, 87, 88, 99, 329, 330, 368 recharge, 96-97 gypsum, 48,73, 76, 79,81, 82,85,86, 97,127,147,148,179,314 (pumping station), 42,53,84 rainfall, 60 H-2 (pumping station), 42, 53 rainfall, 59 H~3 (pumping station), 42, 53 rainfall, 60 Habbaniya, 42, 53, 71,252, 254 air humidity, 64 rainfall, 59 snowfall, 63 Habbaniya Lake, 84, 176 Habbariya,42, 94,96,103 Haditha,42, 53, 77,84, 101, 116 rainfall, 59 Hadramaut. 110, 111 Halocnemum strobilaceum, 23 Halogetonalopecuroi"des, 141, 143,151, 226,248,277,315,315,317,319,321, H~1
324,325,326
373 halophytic vegetation, 19, 23, 82, 178 see vegetation types Haloxylon ammodendron, 36, 151, 226, 276,341 Haloxylon articulatum, 24, 36,143, 156, 160, 186, 187, 188,190,227,229, 233, 234,235,235,236,248,277,294,301, 304,309,327,338,339 Haloxylon persicum, 276 Haloxylon salicornicum, 24, 32, 33, 34, 36,83, 104, 140,141, 143, 147, 150, 151,153,155, 155,156,157,157,158, 162, 165~214 (see specification below), 301, 304,315, 315,328,331,335,338, 338,341,343,373,390 adaptive characteristics, establishment, 211-212 maintenance, 212-213 physiological, 212 reproduction, 209 ~ 21 chemical analysis and nutritive value,
Halogeton~poisening,
232~243
descnption of the species, 165~169 distribution edaphic conditions, 176~181, 177, 180 general, 171~174 Iraq, 174~ 176,175 estimating biomass and production, 378-384 forage preference, 221, 222, 223, 224, 225,229,229 forage production, 252, 252, 253,254, 260,316, 317~319,318 forage value, 227 fuel cutting, 271, 272, 273, 276, 277, 278-289,319,336,337 germination, 185~189 hydration and dehydration, 188, 188 influence of seedwing removal, 186 light conditions, 187 optimal temperature, 186, 187 seed size, 188 rate, 185 moisture content, 244, 245~248, 246, 248 phenology, 181~182, 181 protection, see Khedr~el-Mai exclosure root development (root system), 190, 193,212,213 seed harvest methods, 189 seed, insect damage to, 184 seed morphology, 185 441
seed production and dispersal, 183 seed viability and longevity, 185 seedling development and establishment, 189,190,191,192,211 shoot development, 190,191,192,194 Haloxylon salicomicum - Artemisia herba· alba type, 157-158 Haloxylon salicomicum - Artemisia sco· paria type, 157 Haloxylon salicomicum - Astragalus spi· nosus type, 156-157 Haloxylon salicomicum - Halogeton alopecuroiiJes type, 151 Haloxylon schmittiana, 173 Haloxylon schweinfurthii, 212 hammada, 75, 76, 82, 133 Hammada elegans, 169 Hammada salicomica, 169 Haplophyllum sp.,315, 315 H. tuberculatum. 143 hare, 103 Hatra,42, 123,150,296,297,298, 300, 302
Hatra exciosure, 300,313, 314,330-331 Hejaz,391 Helianthemum, 229, 327 H. ledifolium. 250 H.lippii, 141, 143,148,151,152,153, 154,248, 315,315,321,322,324,325 H. salicifolium, 32,250 Heliotropium bacciferum, 143, 315,321, Heliotropium persicum, 315,321 Heliotropium ramosissimum, 236, 321 'Hema'-system, 371, 391 Herbarium, College of Agriculture (Abu Ghraib), 30, 175,182,197 College of Science (Baghdad), 30, 175 Faculty of Pharmacy (Tehran Univer· sity), 174 Geneva, 30 Kew,30,31,175,182 Leyden, 30 National - of Iraq (Abu Ghraib), 30, 31, 36,43,175,182 Paris, 30 Rustam, 30, 31 University of Basra, 30 University of Mosul, 30 Vienna, 30 Hieracium, 300 Hit,42, 87,116 rainfall, 59 Hordeum, 23 H. glaucum, 327 442
H. pratense, 300 H. sativum, 295 horses, 99,105,106, 107,111,353 houbara, 104 humidity, 63-64 hunting, 100, 104, 109,370 Husaiba, 42, 121,145 Hyaena hyaena L., 102, 117 hyena, 102, 117,389 hygrophilous vegetation, 21, 22-23 Hypecoum pendulum, 250
importance value, 135-136 'increasers'. 133 indicator value of vegetation, 23, 24 see under the separate species Indo-Pakistan, 11, 270 industrial value of shrubs, 14, 19 Institute for Applied Research on Natural Resources, 6, 230 'invaders', 13 3 Iran (Persia), 15, 112,151,171,173,174 Iraq (general), area, 15 boundaries, 15 climate, 17 environment, 14-17 general description, 14 - 25 land use, 18-19,19 physiographic regions, 16 population, 17-18 position in South West Asia, 15 vegetation, 19-25 Iris sisyrinchium, 154 irrigated agriculture, 18, 86, 293, 370 Iskandarya Desert, 33 Jabal .... , see Jebel ... jackal (Asiatic), 102 loculus spp., 103 'jalta',88 jarboa, 103 Jarmo, 109 Jebel Anaiza (Anazeh), 67, 70, 71, 108 Jebel Hamrin (Hamrin Mountains), 73, 111,112,276 Jebel Makhel (Makhul), 73,299 Jebel Sanam, 121, 147, 150, 152 Jebel Shammar, 391 Jebel Sinjar, 73, 297,299 Jeddah.173 Jezira Plains, 68,71,73,73-74 Joe Liban, 152,153, 201 Jordan, 15, 15,67,70,71,145,173,174, 178,180,304,353
Judaidat Arar (Judaidat at Al'ar), 42, 90, 94, 116,155, 174, 176 Jumaima, 42, 116 jungle cat, 102 K-3 (pumping station), 102 Kalahari Desert, 366, 389 Kasim (of Nejd), 391 Kerba1a, 23, 42, 53, 82, 104, 108, 116, 175,179,252,253,254 rainfall,59, 60,62 Kesrah, 101, 102 Khabrat ibu Ghamain, 94 Khedr-el-Mai, 35,42, 116,123,151,152, 176, 179,180,256,25~ 331 Khedr-el-Mai exclosure (fenced area), 122, 212,213,250,255,255,313,314-326,
335,336,341 Kirkuk,30 Koeleria phleoi"des, 32 Kufeh,111 Kuwait, 15, 15,37,67,68,97,116,147, 151,152, 155, 156,169,173,174,178, 179,209,238,239,259,260,265,270, 274
Lagonychium farctum (= Prosopis jizrcta, =Prosopis stephaniana), 23,24,32,143, 161,248, 301,302,328,329,330,331, 331 land evaluation, 363-375, 365, 367, 392 land qualities, 366,367, 368,371-375 LANDSAT-satellite, 297, 299,376 land use planning, 363, 364-375,365, 367 land utilization types, 364, 366,367, 368371,375,377 Lasiurus hirsutus, 100,228 Lebanon, 353 Lepus, L. arabicus, 103 L. capensis, 103 L. europeaus, 103 Lesser Zab (river), 299 Libya, 171, 173,270 Libyan Desert, 125, 173 lime, 79,81, 82,85,127,147,148,314 limestone, 17,70,71,73,74,75,82,85, 96,97,99,147,148,178,179,328 lion (Asiatic), 100, 102. 389 livestock population, 105, 106, 107-108, 352,358,390-391 Lower Wadian (Plain), 68, 70,71,73,82, 84 Lusuf,42 Lycium barbarum, 32,143
Lycium depressum, 274,275 Machmi,94 magnesium (in forage), 231, 232, 235, 236 Malva, 23,222,229 M parviflora, 327,342 mammals, 99 marls, 70, 71, 85, 147, 178 Matthiola oxyceras, 250 Matricaria, 23,229 M. aurea, 250 meat consumption, 354 meat prices, 354 meat production, 354-355,355 Medicago, 21,23,229 medicinal value, 14,350 Medina (of the Hejaz), 391 Mesopotamian Plain (Lower-), Mesopotamia, 16, 16,17,18,23,24,30,31,32 34, 68, 73, 76, 82, 84, 87, 100, 109, 112,175,275 Mia sittien, 42, 158,184 minerals (in forage), see ash minimal area, 125, 127 moisture content of vegetation, 243-249, 244,246,247,248 Moltkiopsis ciliata, 143, 155 monitoring, 364,365, 366,369,375,376, 378,386 Mosul, 15, 18,30,32,42, 53,54,87,296, 300,302
air humidity, 64 air temperature, 55,56 evaporation, 66 radiation, 55 rainfall,59, 60, 63 sunshine. 55 mountainous area (in Iraq), 17,18 see Zagros mountains Mugr an Naam, 42, 102, 116 mules, 105,106,353 Muscat, 173 Najd,113 Najaf,42, 53,73,82,116,254, 276 Najaf (saline) depression, 23, 34, 84, 121 Nasiriyah, 35,42,53,87,103,116,150, 176,178,275 air humidity, 64 air temperature, 55,56 evaporation, 66 radiation, 55 rainfall, 59,60 snowfall, 63 443
natural resources (definition), 349 Negev, 145 Nejd, (=Arabia), 111,391 Nerium oleander, 22 Neurada procumbens, 211 Neutral zone, 15, 70,81,116,153, 176 Nitraria retusa, 23, 104, 235,236, 237, 240,277 Noea mucronata, 143, 159,227,248 nomadism (nomadic pastoralism), see Bedouin(s) North Africa, 11, 14, 37, 170, 173, 174, 259,265,269,270,378 North America, II Nukhaib, 36,42, 43, 53,83, 87, 90, 102, 116, 123, 150, 151, 155, 156, 158, 161, 167, 176,180, 193. 222, 224,254,271, 272,279,279,309,314 rainfall,59, 59 watering points, 93, 94,96 numerical processing, 134, 140 nutritive value, 14,230-243,372,374 see chemical composition oasis, 88, 113 oil,14,87,116,118,357,392 Oman, 110, 173 onager,389 Onobrychis, 21 organic matter (in soils), 76, 79, 81, 82, 85,296 Ornothogalum ulopnyllum, 32 oryx (Arabian oryx), 100, 389 Oryx leucoryx Pallas, 100 ostrich, 104, 117,389 Otis tarda L., 104, 117 overgrazing, 274, 331, 335-344 see grazing pressure Pakistan, 171, 174,364,375 Panicum turgidum, 228 Panthera leo persica Meyer, 102 Paronychia arabica, 315 pastoral nomadism, see Bedouin(s) Peganum harmala. 24,73,141,143, 147, 148,160,221,229,248,300,301,301, 302,302, 303, 328,331, 331,338,339, 373 Penjwin,58 PersiaJee Iran petroCaiClC nonzon, 82 pH (of soils), 48, 81, 127 phenology (of key species), 181, 181-183 Phoenix dactyli[era, 22 phosphorus (in forage), 231, 236, 237, 238,240 444
Phragmites communis (=P. australis), 22 phryganosphere, 340 physiognomy (in vegetation classification),
131
physiography (physiographic units), 15-17, 1~67~4,77,82,145
Picris babylonica, 250 piosphere, 340 Pistacia, 304 Plantago, spp., 154, 222, 229, 250, 327, 342 P. Boissieri, 250 P. ciliata, 250 P. ovata, 250 plant introduction, 7 Platanus orientalis. 22 ploughing (for dryland cultivation), 127, 148,149, 161,293-310,335-344 Central and Upper Jezira, 296-305 general process, 294-296 Lower Jezira, Western and Southern Deserts, 305-310 overall effect, 310 pluvial periods, 108, 117 Poa bulbosa, 24, 32, 222, 228, 304, 330, 331 Poa sinai'ca, 36,159,161,228, 327 population (human), Baghdad,18 distribution, 18 increase, 18,354 Iraq, 17, 18, 110,115 nomadic, 110-ll5, 115, 116 Populus euphratica, 22 Populus nigra, 22 potassium (in forage), 231,232, 233,235, 236 Poterium spinosum, 255 precipitation, see rainfall predators, 14, 100, 103, ll7, 118, 373 primary production, see forage production proper use factor, 372, 374 Prosopis [areta, see Lagonychium [arctum protein (crude protein, D.C.P.), 230,232, 233, 234, 235, 236, 237,238, 239, 239240,241-242, 372 proximate analysis (of forage), 230-232 Prunus, 304 pseudosands,76 Pterocles spp., 104
Qa'ara depression, 67,71,96,108 quartz, 79 Quercus aegilops, 21 Q. calliprinos, 304
Q. infectoria, 21 Q. libani, 21 raids (raiding), 113, 118 rainfall (precipitation), amount, 58-59, 61 changes, III Iraq (general), 17 minimum for cultivation, 293-294, 293 seasonal distribution, 59-63 rainfed agriculture, 18,293,296,297,370, 391 Ramadi,42, 73,94,100,101,116,151, 158,175,176,191,192,193 Ram1a,94 range (land) definition, 7 range resources, 349-352 range site, 378 Ranunculus asiaticus, 32, 161. 300 Rawa,42, 100,116 Razzaza, 108 recharge, see groundwater reconnaissance survey, 7, 258, 365, 366, 376-377 recreation, 349, 350, 373 reg, 75,75,82 relative importance percentage (RIMPvalue), 135-136, 137,141,143,162 reptiles, 103-105 Reseda muricata, 143 reSl11ence (of vegetation), 372-373 resource value of rangelands, 349-352 resource value of shrubs, 12-14 Rhanterium epapposum. 9, 24,32,34,36, 46,69,79,81,140,141,143,147,148, 150,151,152, 152,153,153,154,154, 162, 165-214 (see specification below), 255,265,341 adaptive characteristics establishment, 211, 212 maintenance, 212, 213 physiological, 212 reproduction, 209, 210, 211 chemical analysis and nutritive value, 232-243 description, 169-170 distribution, edapIJic conditions, 177, 179-180 general, 171-174 Iraq, 174-176,175 estimating biomass and production, 378384 forage production, 251, 253, 260 forage value, 226 fuel cutting, 276, 277, 285-286, 285,
287 germination, description, 196, 197,198 rate and temperature, 198,199 moisture content, 245-248,247,248 phenology, 181-182,181 rootsystem,201, 203,213 seed longevity, 200 seed production and dispersal, 195, 195, 210 seedling development and establishment, 200, 201, 202, 211 Rhanterium epapposum - ,--yperus conglomeratus type, 151-152 Rhanterium sua volens, 170,260 Riyadh, 174 rodents, 103, 183 see under species name root (rootsystem), 12, 211,212-213,301, 330 Artemisia herba-alba, 207 Haloxylon salicornicum, 190,193 Rhanterium epapposum, 201-202,201 Rosa can ina, 22 ruderal vegetation, 21, 23-24 Rumaitha, 165 runoff (water), 18, 24,75,96, 145, 159, 176,180,183,274,294,305,310,315, 340,344,370 Rutba, 36, 42, 53, 54, 71, 87, 94, 102, 108, 116, 123, 150,156,158,159,167, 176,180, 191, 192,193, 198,206,252, 254,263,264,27l air humidity, 63, 64 air temperature, 55,56 evaporation, 66 rainfall, 59,59, 60, 63 snowfall,63 soil temperature, 57 watering points, wells, 90, 94, 96 Rutba exc10sure (fenced area), 122, 160, 160,213,222,307,313,314,326-328, 341,383,385,385 Safwan, 79, 176,233 Sahel (Sahelian countries), 340, 391 salinization (salinity), 18,23,84,146, 147 Salix, 22 Sa/sola spp., 126,141,143,147,158,248, 277,302,331 S. baryosma, 143 S. crassa, 34 S. rigida, 36, 222, 223, 224, 225, 227, 229,236,341 S. subaphylla, 248 445
S. vermiculilta, 36,143, 187,227 Samah, 9,42,81,116,123,154,195,198, 200, 201, 247, 250,251,255,256,257, 275,285,28~ 383,384 Samarra,42, 74,112,276 Samawa (Samaweh), 42,71,111,116 rainfall, 59 sampling of vegetation, 44, 121 habitat characteristics, 127 information recorded, 126 precision, 127-130 sample plots, 124-126 selection of stands, 122-123 sanddunes (sanddune area), 71-73, 72, 74, 76,81,82,85,142,147,150-151 sandgrouse, 104 sandstone, 70, 85, 99,147,178 Saudi Arabia, 15, 15,54,67,70,71,79, 89, 92, 102, 111, 116, 155, 156, 169, 170,179,195,260,270,294,353,371, 378 Savignya parvijlora, 250 Schanginia aegyptiaca, 143, 150 Schimpera arabica, 250 Schismus arabicus, 250 Schismus barbatus, 250 Scrophulilria deserti, 143, 302 Scrophulilria xanthoglossa, 330,331 scrub, 12 seed, 301, 325, 330 see Artemisia herba-alba see Haloxylon salicomicum see Rhanterium epapposum seedling development see Artemisia herba-alba see Haloxylon salicomicum see Rhanterium epapposum segetal vegetation, 21, 23-24 Seidlitzia rosmarinus, 23,34,178 selectivity (in grazing), 219-229 Shabicha (Ash Shabicha), 36,42, 53, 70, 90,102,116,271,274 .rainfalL 60 Shabicha tlxclosure, 313, 314,328-330 Shamiya,391 Shanidar cave, 109 Shammar (tribe), 112,296,300 Shatt al Arab, 112 sheep, 23,99, 105, 107, 108, 109, 112, 118,370 contribution to national income, 352-359 see grazing Shanifiya,276 Shinana,94 Shitatha (Shitathe), 23,42, 88, 108,269, 446
275 siltstone, 71 Sinjar (Balad Sinjar), 42, 53, 100,296,298 rainfall, 59, 60 snakes, 104 snowfall, 63 socio-ecological (species) groups, 140,143 sodium (in forage), 231,232,233,235,236 soils chemical analysis, 48,85,147 classification, 77 in the study area, 74-86 Jezira, 85-86 maps, 76-77,80, 366 protection, 14 sampling, 4748, 127 Southern Desert, 79-82 suitability for agricultural use, 18, 19, 20, 76 taxonomic units, 78 Western Desert, 82-85 soil texture, 142-145,144,177, 178,179,
180,180,314
see soils solar radiation, 54-55 South Africa, 125, 133 South America, 11 South West Asia, 5, 11, 14,15,29,259, 269,392 Spain, 174 spiny tailed lizard, 104, 179 springs. 87-88, 94 steppe, 11,12,21,24,31,65,67,110, 174,304,351 Stipa,229 S. capensis, 250 Stipagrostis, 24,143 S. plumosa, 24,100,125,150,151,152, 153,228,248,315,321,322 Struthio camelus, L., 104, 117 stubble grazing, 23 Suaeda spp., 178 S. baccata, 23 S. vermiculilta, 23,178 Sulaimaniya, 22 Sunaisala (depression), 74 sunshine, 54-55 supplementary feeding, 14,265 supplementary irrigation, 18,293,370 Suq as Shuq, 103 surface sealing, 296, 307,340,344 surface water, 88-89,89, 95,375,389 Syria, 15, 15,30,37,70,71,73,84,109, 174,259,270,273,310,353,371 Syrian Desert, 15,32,161,165
Syrian wild ass, 100 survey, 7,41,76-77, 122, 258,364-378, 392
see reconnaissance survey Takhadid,42 Tammam,111 Tamarix, 22,97,100,141,143,147,161, 274,275 T. aphylla, 275 T. articulata, 32 Tamarix sp. - Aeluropus lagopoides type, 161 Tar Kshaife, 108 Taurus, 110 Tel Afar, 42,53,296,297,298 Tel Lachm, 150, 178,276 temperature, air -, in the study area, 55-57,64 data collection, 48 Iraq (general), 17 soil-, in the study area, 57-58 Tethys Sea, 17 Teucrium oliverianum, 36 Teucrium polium, 36,143,227,236,237, 302 'thaqab',88 Tharthar (Wadi Tharthar, Tharthar depression, Lake Tharthar), 73,74,85,97,101, 151,175,276,389 thorn-cushion vegetation, 21 Tigris (river), River Tigris, 15, 16, 17, 24, 30,74,97,110,111,276,296,299 Tikrit,42 toxicity of forage (poisenous plants), 373 Traifawi,42, 116,178,302,303 Tribulus terrestris, 327 Trifolium, 21,23,229 Trigonella, 23,229 T. stellata, 250 Triticum durum, 295 Tunisia, 259, 260,270,273 Tuquiyid, 70 Turkey, 15,15,171,174 Typha angustifolia, 22 Ukhaidir (area), 34, 42, 108, 121, 123, 158,179,184,252,254,382,383 air humidity, 63, 64 air temperature, 64 soil temperature, 58 Urn Quasr, 151,152, 152,169,233 Umm Rahal, 73 Upper Wadian (Plain), 68, 70-71, 82 uprooting, 286-288, 294
see fuel t cutting) Ur, 32, 42, 67, 73, 79, 81, 151, 276,277 urbanization, 114, 116 Uromastix aegyptius L., 104,179 USA (United States of America), 12, 35, 231,371,373,378,391 USSR, 174, 378 utilization (of the rangelands), 219-344 see fuel (cutting) see grazing see ploughing Varanus griseus Daudin, 104 vegetation types, 41, 121-162 classification 130-142 description, 142-162 (seperately given under the species) sampling for -, 122-130 Vicia, 23 Vitis vinifera, 22 Vulpes rilppelli sabaena Pocock, 102 Vulpes vulpes L., 102 Vulpes vulpes arabica Thomas, 102
Wadi AI-Auja, 94 Wadi AI-Khir, 70, 82 Wadi AI-Muhammadi, 94 Wadi AI-Walej, 71,94 Wadi Batin (AI-Batin, Batn), 70, 71, 94, 97,370,391 Wadi Ghadef, 70 Wadi Hamdh, 178 Wadi Hauran, 70, 71, 108 Wadi Rumh, 391 Wadi Sidr, 275 Wadi Tibel mbal), 224, 308 Wadi Ubaiyidh, 10; 71, 83, 157, 193, 276, 279,28~306, 309,338 Wadian (Plain), 94 see Upper Wadian Plain see Lower Wadian Plain watering points, see wells see surface water water quality, 90, 96, 97-99, 98,373 water resources, 87-99 weeds, 23, 125,330 wells, 87, 89-94, 95, 96, 97, 113, 116, 118,270,276,284,301,329,338,340, 344,368,391 drilled, 90, 91-94, 93,95,96 hand~ug,89,91,92,93,94,95,96
in land evaluation, 373 wolf, 102 Yemen, 110, 111,391 447
Zagros mountains, 16, 16,17,58,110 Zarga area, 103 Zilla spinoSQ, 36, 143, 156, 158, 160, 178, 179,221,229,248,307,310,328,338 Zizyphus nummularia, 32,33,36,88,221, 274,275, 275 zoochory, 183, 197
448
Zubarr, 36,42, 79,97,116,156,271,275 Zygophyllum coccineum, 34, 141, 143, 147, 148, 156, 235, 236, 237, 240,248 Zygophyllum coccineum - Cornulaca leucacantha type, 156 Zygophyllum dumosum, 255 Zygophyllum propinquum, 147, 156